etau—these two, namely Rama and Krsna; hi—certainly; visvasya—of the universe; ca—and; bija-yoni—both the cause and ingredient; ramah—Balarama; mukundah—Krsna; purusah—the original Maha-Visnu; pradhanam—material energy; anviya—after entering; bhutesu—into the material elements; vilaksanasya—of varieties of manifestation; jnanasya—of knowledge; ca—also; isate—are the controlling power; imau—both of Them; puranau—are the original cause.